Russia have lost the Olympic women's 4x400-metre silver medal from 2008 due to the re-analysis of Anastasia Kapachinskaya's doping sample testing positive, just two days after having the women's 4x100m gold withdrawn.

Kapachinskaya tested positive for forbidden substances, along with two other Russians from different sports, in re-tests of samples, the International Olympic Committee (IOC) said Friday.

Her sample returned positive for the prohibited stanozolol and dehydrochlormethyltestosterone (turinabol). Russia finished second in Beijing behind the United States and their disqualification promotes Jamaica to silver and Belarus into bronze.

The 36-year-old Kapachinskaya has also had her results removed from the 400m, where she finished fifth.

On Wednesday Russia lost the 4x100-metre relay gold from Beijing after Yulia Chermoshanskaya tested positive for the same banned substances as Kapachinskaya.

The IOC has conducted many re-tests in recent months and said that 98 athletes had positive results from Beijing and London 2012. The IOC stores samples for 10 years and re-tests with the latest scientific methods.

Russia is currently accused of running a widespread and state-sponsored doping scheme. Its entire athletics team, apart from US-based long-jumper Darya Klishina, was suspended for the current Rio Games.

The others who had their positive tests announced Friday were Alexander Pogorelov, who finished fourth in the decathlon, and Ivan Yushkov, who finished 10th in the shot put.

Pogorelov, whose biggest achievement was bronze at the 2009 Berlin world championships, tested positive for turinabol. Yushkov, who has never won a major medal, tested positive for stanozolol, turinabol and oxandrolone.

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